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Emi Buendia scored with the last kick of the game as Aston Villa stunned Arsenal to move three points behind the Premier League leaders.
Leandro Trossard's sixth goal of the season looked to have earned a point for Arsenal after he cancelled out Matty Cash's opener.
But Buendia struck in the 95th minute as Unai Emery's side moved into second in the table.
Arsenal, who blew Bayern Munich and Tottenham away last month, suffered their first defeat in the league since losing 1-0 at Liverpool in August.
Villa Park erupted after substitute Buendia scored to give Villa their seventh straight game in all competitions. They have won nine of their past 10 in the league.
Arsenal's Eberechi Eze had a goal disallowed for offside before the hosts took a first-half lead when Arsenal - especially Eze - switched off to allow Cash to drill in at the far post.
Arsenal emerged re-energised after the break when Mikel Arteta introduced Viktor Gyokeres and Trossard, who levelled with his 50th Premier League goal, having already gone close.
Emi Martinez brilliantly tipped Martin Odegaard's shot over as Arsenal looked for a winner, but Buendia stole the points in dramatic style, curling in after Arsenal failed to clear.
Arsenal's match against Villa was a game that felt like it went under the radar because of the run of matches the Gunners had before this one.
And the loss to Emery's side will only highlight the problems that, despite Arsenal's excellent start to the season, have been mounting.
They have battled injuries to key players throughout the season and this game was no different with the news that Cristhian Mosquera had been ruled out for about six weeks before the match.
That meant Jurrien Timber had to play at centre-back, with Ben White coming in at right-back, which is how the Arsenal defence ended up during the win over Brentford in midweek.
White starting at right-back and captain Odegaard returning in midfield meant that White, Odegaard and Bukayo Saka were reunited on the right side of Arsenal's team.
The relationship between the three has been key to Arsenal's attack in recent seasons and the majority of the Arsenal play during the match went down that side.
But Odegaard coming into the team in the middle meant that Eze was shifted to the left-wing, but in that position there are more defensive responsibilities to carry out, and the England international switched off for Cash's goal.
Mikel Merino was removed at half-time alongside Eze, and Trossard came on to score for his sixth goal of the season. He continues to be one of Arsenal's standout players of the season.
The match was more open than Arteta would have liked and Arsenal missed the calmness of William Saliba and Gabriel and their presence in both boxes.
Saturday's match was only the fourth time since the start of the 2022-23 season that Arsenal started a league game without Saliba or Gabriel.
Buendia's winner would likely have been dealt with better if the two centre-backs had been on the pitch, and, with seven more games this month, Arteta will be hoping this is just a blip and not the start of a considerable drop in form. (BBC)